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The rump and upper tail-coverts are delicately patterned with dark vermiculations and fine wavy barring, the extent of which varies with subspecies.

The underwing coverts and undertail coverts are similar but tend to be more strongly barred in brownish-black. The primaries and secondaries are brown with broad dark brown bars and dark brown tips, and grey or buff irregular lines.

A complete moult takes place each year between July and December. The chin and throat are white with a brownish central streak. The feathers of the upper breast generally have brownish-black centres and reddish-brown edges except for the central ones which have white edges.

The chin and throat may appear white continuing down the center of the upper breast. The lower breast and belly feathers are creamy-brown to tawny buff to off-white with a variable amount of fine dark wavy barring, on a tawny-buff ground colour.

The legs and feet which are feathered almost to the talons are likewise marked on a buff ground colour but more faintly.

The tail is tawny-buff, mottled dark grey-brown with about six black-brown bars. The bill and feet are black. The iris is most often orange but is fairly variable.

In some European birds, the iris is a bright reddish, blood-orange colour but then in subspecies found in arid, desert-like habitats, the iris can range into an orange-yellow colour most closely related species generally have yellowish irises, excluding the Indian eagle-owl.

Thus they have quite high wing loading. The great horned owl has even smaller wings 0. The golden eagle has slightly lower wing loading proportionately 0.

Some other owls, such as barn owls, short-eared owls Asio flammeus and even the related snowy owls have lower wing loading relative to their size and so are presumably able to fly faster, with more agility and for more extended periods than the Eurasian eagle-owl.

Accipitrids use their talons to inflict organ damage and blood loss, whereas typical owls use their feet to constrict their prey to death, the talons serving only to hold the prey in place or provide incidental damage.

The talons of the Eurasian eagle-owl are very large and not often exceeded in size by diurnal raptors.

Unlike the great horned owls, the overall foot size and strength of the Eurasian eagle-owl is not known to have been tested, but the considerably smaller horned owl has one of the strongest grips ever measured in a bird.

The feathers of the ear tufts in Spanish birds when not damaged were found to measure from In the female, the ear opening averages Examples of owls with more complicated ear structures and deeper facial disc are barn owls, long-eared owls Asio otus and boreal owls Aegolius funereus.

Given the uncomplicated structure of their ear openings and relatively shallow, undefined facial disc, hunting by ear is secondary to hunting by sight in the eagle-owl, this seems to be true for Bubo in general.

It is likely that more sound-based hunters such as the aforementioned species focus their hunting activity in more complete darkness.

Also owls with white throat patches such as the Eurasian eagle-owl are more likely to be active in low light conditions in the hours before and after sunrise and sunset rather than the darkest times in the middle of the night.

The boreal and barn owls, to extend these examples, lack obvious visual cues such as white throat patches puffed up in displaying eagle-owls , again indicative of primary activity being in darker periods.

The great size, bulky, barrel-shaped build, erect ear tufts and orange eyes render this as a distinctive species. Nevertheless, the latter is still distinctively an ear-tufted Eurasian eagle-owl and lacks the pure white background colour and variable black spotting of the slightly smaller species which has relatively tiny, vestigial ear tufts that have only been observed to have flared on rare occasions.

Long-eared owls in Eurasia have vertical striping like that of the Eurasian eagle-owl, while long-eared owls in North America show a more horizontal striping like that of great horned owls.

Whether these are examples of mimicry either way is unclear but it is known that both Bubo owls are serious predators of long-eared owls.

The same discrepancy in underside streaking has also been noted in the Eurasian and American representations of the grey owl.

Three fish owls appear to overlap in range, the brown Ketupa zeylonensis in at least northern Pakistan , probably Kashmir and discontinuously in southern Turkey, the tawny K.

Fish owls are distinctively different looking, possessing more scraggy ear tufts that hang to the side rather than sit erect on top of the head, generally have more uniform, brownish plumages without the contrasting darker streaking of an eagle-owl.

Fish owls, being tied to the edges of freshwater where they hunt mainly fish and crabs, also have slightly differing, and more narrow, habitat preferences.

In the lower Himalayas of northern Pakistan and Jammu and Kashmir, along with the brown fish owl, the Eurasian eagle-owl at the limit of its distribution may co-exist with at least two to three other eagle-owls.

The dusky is usually found in slightly more enclosed woodland areas than Eurasian eagle-owls. Far to the west, the Pharaoh eagle-owl B. Although also relatively similar to the Eurasian eagle-owl, the Pharaoh eagle-owl is distinguished by its smaller size, paler, more washed-out plumage and the diminished size of its ear-tufts.

In the Eurasian eagle-owl, this happens in stages and the first moult starts the year after hatching with some body feathers and wing coverts being replaced.

The next year the three central secondaries on each wing and three middle tail feathers are shed and regrow, and the following year two or three primaries and their coverts are lost.

In the final year of this post-juvenile moult, the remaining primaries are moulted and all the juvenile feathers will have been replaced.

Another moult takes place during years six to twelve of the bird's life. This happens between June and October after the conclusion of the breeding season and again it is a staged process with six to nine main flight feathers being replaced each year.

Such a moulting pattern lasting several years is repeated throughout the bird's life. The Eurasian eagle-owl is a member of the genus Bubo , which may include either 22 or 25 extant species.

Almost all the larger owl species in the world today are included in Bubo. Based on an extensive fossil record and a central distribution of extant species on that continent, the Bubo appears to have evolved into existence in Africa, although early radiations seem to branch from southern Asia as well.

A few of the other species of Bubo seem to have been derived from the Eurasian eagle-owl, making it a " paraspecies ", or they at least share a relatively recent common ancestor.

The Pharaoh eagle-owl, distributed in the Arabian Peninsula and sections of the Sahara Desert through North Africa where rocky outcrops are found, was until recently considered a subspecies of the Eurasian eagle-owl.

It appears that the Pharaoh eagle-owl differs about 3. It appears to have separated from other Bubo at least 4 million years ago. The fourth and most famous derivation of the evolutionary line that includes the Eurasian eagle-owl is the great horned owl , which appears to have been the result of primitive eagle-owls spreading into North America.

Somehow, genetic materials indicate the spotted eagle-owl appears to share a more recent ancestor with the Indian eagle-owl than with the Eurasian eagle-owl or even the sympatric Cape eagle-owl.

Fossils from southern France have indicated that during the Middle Pleistocene , Eurasian eagle-owls this paleosubspecies is given the name B.

Eagle-owls are distributed somewhat sparsely but can potentially inhabit a wide range of habitats, with a partiality for irregular topography.

Grassland areas such as alpine meadows or desert -like steppe can also host them so long as they have the cover and protection of rocky areas.

Also, they may be attracted to the vicinity of riparian or wetlands areas, due to the fact that the soft soil of wet areas is conducive to burrowing by the small, terrestrial mammals normally preferred in the diet, such as voles and rabbits.

Pine and other coniferous stands are often preferred in great horned owls as well due to the constant density, which make it more likely to overlook the large birds.

Although found in the largest numbers in areas sparsely populated by humans, farmland is sometimes inhabited and they even have been observed living in park-like or other quiet settings within European cities.

The number is expected to increase due to the growth of the European rabbit population in Helsinki. European hares Lepus europaeus , the often preferred prey species by biomass of the eagle-owls in their natural habitat, live only in rural areas of Finland, not in the city centre.

The match was interrupted for six minutes. The owl was named "Helsinki Citizen of the Year" in December The Eurasian eagle-owl is largely nocturnal in activity, as are most owl species, with its activity focused in the first few hours after sunset and the last few hours before sunrise.

In such areas, full nightfall is essentially non-existent at the peak of summer, so eagle-owls must presumably hunt and actively brood at the nest during daylight.

These are dotted along the outer edges of the eagle-owl's territory and they are visited often but only for a few minutes at a time.

Vocal activity is almost entirely confined to the colder months from late fall through winter, with vocal activity in October through December mainly having territorial purposes and from January to February being primarily oriented towards courtship and mating purposes.

The widely used name in Germany as well as some other sections of Europe for this species is uhu due to its song. The Eurasian eagle-owl rarely assumes the so-called "tall-thin position", which is when an owl adopts an upright stance with plumage closely compressed and may stand tightly beside a tree trunk.

Among others, the long-eared owl is among the most often reported to sit with this pose. It has, unusually for an owl, also been known to soar on updrafts on rare occasions.

The latter method of flight has led them to be mistaken for Buteos , which are smaller and quite differently proportioned.

Even those near the northern limits of their range, where winters are harsh and likely to bear little in food, the eagle-owl does not leave its native range.

There are cases from Russia of Eurasian eagle-owls moving south for the winter, as the icebound , infamously harsh climate there is too severe even for these hardy birds and their prey.

Similarly, Eurasian eagle-owls living in the Tibetan highlands and Himalayas may in some cases vacate their normal territories when winter hits and move south.

Even in those two examples, there is no evidence of consistent, annual migration by Eurasian eagle-owls and the birds may eke out a living on their normal territories even in the sparsest times.

Eurasian eagle-owls are strictly territorial and will defend their territories from interloping eagle-owls year around, but territorial calling appears to peak around October to early January.

The high prominence of singing perches allows their song to be heard at greater distances and lessens the need for potentially dangerous physical confrontations in the areas where territories may meet.

When taxidermied specimens with flared white throats were placed around the perimeter of eagle-owl territories, male eagle-owls reacted quite strongly and often attacked the stuffed owl, reacting more mildly to a stuffed eagle-owl with a non-flared white throat.

Females were less likely to be aggressive to mounted specimens and did not seem to vary in their response whether exposed to the specimens with or without the puffed up white patch.

There may be mutual bowing, billing and fondling before the female flies to a perch where coitus occurs, usually taking place several times over the course of a few minutes.

The male selects breeding sites and advertises their potential to the female by flying to them and kneading out a small depression if soil is present and making staccato notes and clucking noises.

Several potential sites may be presented, with the female selecting one. Eurasian eagle-owls normally nest on rocks or boulders, most often utilizing cliff ledges and steep slopes, as well as crevices, gullies, holes or caves.

Rocky areas that also prove concealing woodlots as well as, for hunting purposes, that border river valleys and grassy scrubland may be especially attractive.

Among the eagle-owls of the fairly heavily wooded wildlands of Belarus , they more commonly utilize nests built by other birds than most eagle-owls, i.

While it may be assumed that the eagle-owl is too large to utilize tree hollows, when other large species like the great grey owl have never been recorded nesting in one, the even more robust Blakiston's fish owl nests exclusively in cavernous hollows.

In Engadin , Switzerland, the male eagle-owl hunts until the young are 4 to 5 weeks old and the female spends all her time brooding at the nest.

After this point, the female gradually resumes hunting from both herself and the young and thus provides a greater range of food for the young.

The species is often rather less aggressive than some other owls, including related species like the spot-bellied eagle-, great horned and snowy owls, many of the northern Strix species and even some rather smaller owl species, which often fearlessly attack any person found to be nearing their nests.

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After his friend Joo-hwan picks him up from the police station, Dae-su is kidnapped and wakes up in a sealed hotel room, where food is delivered through a trap-door.

Watching TV, Dae-su learns that his wife has been murdered and he is the prime suspect. He passes the time shadowboxing , planning revenge and attempting to dig a tunnel to escape.

Fifteen years pass since he was imprisoned. It is now , a new century. Just before digging himself to freedom, Dae-su is sedated and hypnotized.

He wakes up on a rooftop and sees a man on the ledge ready to jump to his death. The man asks him, "Even though I am no better than a beast, don't I have the right to live?

Afterward, a mysterious beggar gives him money and a cell phone. Dae-su enters a sushi restaurant where he meets Mi-do, a young chef.

He receives a taunting phone call from his captor, collapses, and is taken in by Mi-do. In a frenzy, Dae-su tries to force himself on Mi-Do in the bathroom of her apartment, however she is armed with a knife and successfully fends him off.

Dae-su attempts to flee the apartment, but a sympathetic and intrigued Mi-do confronts him. Mi-do explains to Dae-su that she is romantically interested in him, but that they hardly know each other and she is not ready to be intimate.

They reconcile and begin to form a bond. After he recovers, Dae-su tries to find his daughter and his former prison. Discovering that she was adopted, he gives up trying to contact her.

He locates the Chinese restaurant that made his food and finds the prison by following a deliveryman. It is a private prison, where people can pay to have others incarcerated.

Dae-su tortures the warden, Mr. Park, who reveals that Dae-su was imprisoned for "talking too much". He is then attacked by guards and is stabbed but manages to defeat them.

Dae-su's captor is revealed to be a wealthy man named Lee Woo-jin. Woo-jin gives him an ultimatum: if Dae-su can uncover the motive for his imprisonment within five days, Woo-jin will kill himself.

Otherwise, he will kill Mi-do. Dae-su and Mi-do get close and have sex. Meanwhile, Joo-hwan tries to contact Dae-su with important information but is murdered by Woo-jin.

Dae-su eventually recalls that he and Woo-jin had gone to the same high school, and he had witnessed Woo-jin committing incest with his own sister.

He told Joo-hwan what he had seen, which led to his classmates learning about it. Rumors spread and Woo-jin's sister killed herself, leading a grief-stricken Woo-jin to seek revenge.

In present day, Woo-jin cuts off Mr. Park's hand, causing Mr. Park and his gang to join forces with Dae-su.

Dae-su leaves Mi-do with Mr. Park and sets out to face Woo-jin. At Woo-jin's penthouse , Woo-jin reveals that Mi-do is actually Dae-su's daughter.

Woo-jin had orchestrated everything by using hypnosis to guide Dae-su to the restaurant, arranging for them to meet and fall in love so that Dae-su will experience the same pain of incest that he did.

He reveals that Mr. Park is still working for him and threatens to tell the truth to Mi-do. Dae-su apologizes for his involvement in the death of Woo-jin's sister and humiliates himself by imitating a dog and begging.

When Woo-jin is unimpressed, Dae-su cuts out his own tongue as a sign of penance. Woo-jin finally accepts Dae-su's apology and tells Mr.

Park to hide the truth from Mi-do. He then drops the device he claimed was the remote to his pacemaker and begins walking away.

Dae-su activates the device to kill him, only to find it is actually a remote for loudspeakers, which play an audio recording of Mi-do and Dae-su having sex.

As Dae-su collapses in despair, Woo-jin enters the elevator, recalls his sister's suicide, and shoots himself in the head. Some time later, Dae-su finds the hypnotist to erase his knowledge of Mi-do being his daughter so that they can stay happy together.

To persuade her, he repeats the question he heard from the man on the rooftop, and the hypnotist agrees. Afterward, Mi-do finds Dae-su lying in snow but there are no signs of the hypnotist, leaving it ambiguous if Dae-su really met her.

Mi-do confesses her love for him and the two embrace. Dae-su breaks into a wide smile, which is then slowly replaced by a look of pain.

The corridor fight scene took seventeen takes in three days to perfect and was one continuous take; there was no editing of any sort except for the knife that was stabbed in Oh Dae-su's back, which was computer-generated imagery.

The script originally called for full male frontal nudity, but Yoo Ji-tae changed his mind after the scenes had been shot.

Other computer-generated imagery in the film includes the ant coming out of Dae-su's arm according to the making-of on the DVD the whole arm was CGI and the ants crawling over him afterwards.

The octopus being eaten alive was not computer-generated; four were used during the making of this scene. Actor Choi Min-sik, a Buddhist , said a prayer for each one.

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